The gradient of
water concentration that exists across the cortex creates a pushing
force called "root pressure", i.e. a pressure that "pushes" the water
across. Root pressure can be demonstrated by cutting a stem at soil
level, after a time, droplets of water can be seen exuding from the
cut surface. In some plants the process occurs naturally as "guttation",
when droplets of water are forced through special pores (hydathodes)
on the leaf edges. It is likely that the process of root pressure requires
energy, as metabolic poisons halt the process.
While root pressure
might account for some of the upward movement of water in small herbaceous
plants, it is insufficient to overcome the force of gravity and move
water up tall plants. To determine what else is involved in ensuring
water gets to the sites of photosynthesis, let us look at the plant's
water transporting tissue, the xylem.
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